Abundance: 0.15 to 13.36% (median 4.34%)

Latitudinal range: 41.6° to 49.6°

Habitats: (6) only

Found in 6 samples

France: Caune de l'Arago (Unit D) (Middle Pleistocene), Orgnac 3 (level 1) (Middle Pleistocene)

Italy: Isernia La Pineta (3 colluvio level) (Middle Pleistocene)

Montenegro: Trlica (Bed 10) (Early Pleistocene), Trlica (Bed 5) (Middle Pleistocene)

Ukraine: Medzhibozh 1 (horizon 16a) (Middle Pleistocene)

See also Ursus, Ursus americanus, Ursus arctos, Ursus bibersoni, Ursus boeckhi, Ursus dolinensis, Ursus etruscus, Ursus kudarensis, Ursus maritimus, Ursus minimus, Ursus sp., Ursus spelaeus, Ursus thibetanus, Ursus vitabilis