Abundance: 0.01 to 82.20% (median 3.27%)

Latitudinal range: -37.0° to 51.9°

Habitats: (44) only

Found in 44 samples

Australia: Grant Hall (Layer 2) (Late Pleistocene), Grant Hall (Layer 3) (Late Pleistocene)

Brazil: Lapa do Santo (Early Holocene) (Holocene), Lapa do Santo (Middle Holocene) (Holocene)

Ecuador: La Chimba (Holocene), Challuahamba (Holocene)

Ethiopia: Bodo D'ar (Middle Pleistocene)

France: Saint-Privat-des-Vieux (Early Oligocene)

Greece: Lakonis Cave I (hearth complex) (Late Pleistocene), Lakonis Cave I (upper bone breccia) (Late Pleistocene), Lakonis Cave I (lower bone breccia) (Late Pleistocene)

India: Chandankhera (Period III: Satavahana) (Holocene), Chandankhera (Period IV: Vakataka) (Holocene)

Italy: Rifreddo (Middle Pleistocene), La Sassa Cave (Late Pleistocene), Pirro Nord 13 (Early Pleistocene)

Kazakhstan: Tasbas (Phase 2a) (Holocene)

Kenya: Usenge 3 (Urewe) (Holocene)

Mexico: El Zurdo (Holocene)

Romania: Noviodunum (TPF sector, Middle Ages) (Holocene)

Russia: Kuznetsovka (Middle Pleistocene)

South Africa: Plovers Lake (Late Pleistocene)

Spain: Fuente Nueva 3 (layer 5) (Early Pleistocene), Venta Micena 4-I (Early Pleistocene), Venta Micena 4-II (Early Pleistocene), Barranco Léon (Early Pleistocene)

Tanzania: Ewass Oldupa (Trench 2) (Early Pleistocene), Ewass Oldupa (Trench 3) (Early Pleistocene), Ewass Oldupa (Trench 5) (Early Pleistocene)

Thailand: Ban Rai rockshelter (Holocene), Tham Lod rockshelter (Unit 1) (Late Pleistocene), Tham Lod rockshelter (Unit 2) (Late Pleistocene), Tham Lod rockshelter (Unit 3) (Late Pleistocene)

United States: Vess Cave (Late Pleistocene), McKinley Site (Zone A) (Holocene), McKinley Site (Zone B) (Holocene), Steck (Holocene), Deadman Slough (Excavation Unit) (Holocene), Sucices Site (Excavation Square 2) (Holocene), Watson Brake (3.2 mm fraction) (Holocene), Rattlesnake Point Pueblo (Unit 2) (Holocene), Hale Farm (Holocene), Pine Hill (Holocene), Kincaid Shelter (Late Pleistocene)

See also Testudines