Abundance: 0.04 to 5.57% (median 0.42%)

Latitudinal range: 40.9° to 50.6°

Habitats: (3), Mediterranean woodland (1), temperate broadleaf/mixed forest (2)

Found in 6 samples

Greece: Lake Prespa National Park

Italy: Poggi di Prata

Montenegro: Great Ulcinj Beach

Poland: Zabia Cave (Layer 21) (Early Pleistocene)

Ukraine: Grot Skeliastyi (III-3B) (Late Pleistocene), Grot Skeliastyi (III-1) (Holocene)

See also Natrix, Natrix longivertebrata, Natrix maura, Natrix natrix, Natrix sipedon, Natrix sp.