Abundance: 0.03 to 12.10% (median 0.77%)

Latitudinal range: 37.7° to 50.6°

Habitats: (8), Mediterranean woodland (2), temperate broadleaf/mixed forest (2)

Found in 12 samples

France: Coudoulous I (Faunal Unit III) (Middle Pleistocene), Coudoulous I (Faunal Units IV and V) (Middle Pleistocene), Coudoulous I (Faunal Unit VI) (Middle Pleistocene), Coudoulous I (Faunal Unit VII) (Middle Pleistocene)

Greece: Lake Prespa National Park

Italy: Poggi di Prata, Cava Muracci (Late Pleistocene)

Montenegro: Great Ulcinj Beach

Poland: Zabia Cave (Layer 21) (Early Pleistocene), Zabia Cave (Layer 19) (Early Pleistocene)

Turkey: Madran Mountain

Ukraine: Grot Skeliastyi (III-3-x) (Late Pleistocene)

See also Natrix, Natrix longivertebrata, Natrix maura, Natrix sipedon, Natrix sp., Natrix tessellata