Latitudinal range: 23.0° to 53.0°

Found in 5 samples

China: Mocun Cave (Late Pleistocene)

France: Payre (ensemble F) (Middle Pleistocene), Payre (ensemble D) (Late Pleistocene)

Japan: Matsugae (Middle Pleistocene)

Russia: Imanay Cave (Pleistocene Layer 4) (Late Pleistocene)

See also Cuon, Cuon alpinus