Mass: 156 kg based on Kingsley et al. 1983

Diet: frugivore-carnivore based on Berns and Hensel 1972

Abundance: 0.01 to 36.13% (median 0.81%)

Latitudinal range: 30.3° to 68.8°

Habitats: (53), boreal forest/taiga (1), desert/xeric shrubland (1), montane grassland (4), temperate broadleaf/mixed forest (3), temperate coniferous forest (3), temperate savanna (2), tundra (1)

Found in 68 samples

Canada: Grassy Bay (Holocene)

China: Suojia Township, Yanchiwan National Nature Reserve, Songhua River (Late Pleistocene), Kanas National Nature Reserve

Croatia: Northern Velebit National Park

France: Les Canalettes (Layer 3) (Late Pleistocene), Les Canalettes (Layer 4) (Late Pleistocene), Coudoulous I (Faunal Unit II) (Middle Pleistocene), Coudoulous I (Faunal Unit III) (Middle Pleistocene), Coudoulous I (Faunal Units IV and V) (Middle Pleistocene), Coudoulous I (Faunal Unit VI) (Middle Pleistocene), Romain-la-Roche (Middle Pleistocene), Lazaret (C II supérieur UA 13-17) (Middle Pleistocene), Payre (ensemble G) (Middle Pleistocene), Payre (ensemble F) (Middle Pleistocene), Payre (ensemble D) (Late Pleistocene), Tourville-la-Rivière (level D1) (Middle Pleistocene)

Georgia: Dzudzuana Cave (lower area, Unit B) (Late Pleistocene)

Iran: Lar Protected Area, Wezmeh Cave (Late Pleistocene), Golestan National Park

Israel: Marj Rabba (Holocene)

Italy: Western Trento Province (2012), Caverna degli Orsi (sondage A, Unit 11.2) (Late Pleistocene), Riparo Biarzo (SU 5) (Late Pleistocene), Riparo Biarzo (SU 4 + 3B) (Holocene), Riparo Biarzo (SU 3A) (Holocene), Mondeval de Sora (SU 8) (Holocene), Mondeval de Sora (SU 31) (Holocene), La Sassa Cave (Late Pleistocene), Grotta Cucgiliana (Late Pleistocene)

Lebanon: Moghr el-Ahwal Cave 3 (Phases F & E) (Late Pleistocene)

North Korea: Chongphadae Cave (Late Pleistocene)

Poland: Borsuka Cave (Layer VI) (Late Pleistocene), Borsuka Cave (Layer III) (Holocene)

Portugal: Gruta de Figueira Brava (Area C) (Late Pleistocene)

Romania: Southern Transylvania, Noviodunum (TPF sector, Middle Ages) (Holocene)

Russia: Geographical Society Cave (Late Pleistocene), Chagyrskaya Cave (Layer 5) (Late Pleistocene), Chagyrskaya Cave (Layer 6) (Late Pleistocene), Cheremukhovo-1 (Bed 4) (Holocene), Imanay Cave (Pleistocene Layer 1) (Late Pleistocene), Imanay Cave (Pleistocene Layer 2) (Late Pleistocene), Imanay Cave (Pleistocene Layer 3) (Late Pleistocene), Imanay Cave (Pleistocene Layer 4) (Late Pleistocene), Khayrgas Cave - Layer 6 (1998-1999 pit) (Late Pleistocene), Khayrgas Cave - Layer 7 (1998–1999 pit) (Late Pleistocene), Byzovaya (Late Pleistocene)

Spain: Cueva del Angel (Late Pleistocene), Bolomor Cave (Level IV) (Middle Pleistocene)

Tunisia: Meninx (Holocene)

Turkey: Yaylacik Research Forest, Marmaris, Barhal Stream, Direkli Cave (Holocene)

United Kingdom: Lynford Quarry (Late Pleistocene), Balderton Sand and Gravel (Middle Pleistocene)

United States: Dalton Highway, Welsh Cave (Late Pleistocene), Wewukiyepuh (Holocene), Polecat Creek (alluvial gravel) (Late Pleistocene), Schulze Cave (layer B) (Holocene), Schulze Cave (layer C1) (Late Pleistocene), Molybdenum Ridge (triggered detections), Molybdenum Ridge (time-lapse detections), Black Rapids Training Area (triggered detections)

♀ body mass	105000 kg	N = 45	Kingsley et al. 1983
♀ body mass	198900 kg	N = 19	Kingsley et al. 1983
♀ body mass	91000 kg	N = 24	Kingsley et al. 1983
♀ body mass	162500 kg	N = 18	Kingsley et al. 1983
♂ body mass	190600 kg	N = 41	Kingsley et al. 1983
♂ body mass	258300 kg	N = 10	Kingsley et al. 1983
♂ body mass	190400 kg	N = 24	Kingsley et al. 1983
♂ body mass	222400 kg	N = 19	Kingsley et al. 1983

See also Ursus, Ursus americanus, Ursus bibersoni, Ursus boeckhi, Ursus deningeri, Ursus dolinensis, Ursus etruscus, Ursus kudarensis, Ursus maritimus, Ursus minimus, Ursus sp., Ursus spelaeus, Ursus thibetanus